Steccherinum shangzhouense L. L. Wan, L. S. Zhang, Shun Liu & B. K. Cui 2026, sp. nov.

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Steccherinum shangzhouense L. L. Wan, L. S. Zhang, Shun Liu & B. K. Cui sp. nov. Figs 8, 9 Diagnosis. Steccherinum shangzhouense is characterized by resupinate and membranaceous basidiomata, white to cream, tuberculate to slightly grandinioid and distinctly cracked hymenial surface, monomitic hyphal system with simple-septate generative hyphae, absence of cystidia and skeletocystidia, clavate basidia, and broadly ellipsoid to oblong ellipsoid basidiospores (5.2–7.9 × 2.5–5 µm). Type. • China; Shaanxi Province, Shangluo, Shangzhou District; 34.020372°N, 109.983861°E; elevation 852 m; on fallen angiosperm branch; 23 August 2025; Cui 24845 (holotype, BJFC). Etymology. The epithet “ shangzhouense ” (Lat.): referring to Shangzhou District, Shaanxi Province, where the type specimen was collected. Description. Fruiting body. Basidiomata annual, resupinate, closely adnate to the substrate, membranaceous, soft when fresh, becoming brittle upon drying, white to cream when fresh, turning pale gray to grayish upon drying, frequently developing conspicuous cracks, 13–18 cm long, 1–2 cm wide, and up to 120 µm thick. Hymenial surface tuberculate to slightly grandinioid, composed of irregular granules, white to cream when fresh, becoming pale gray upon drying, distinctly cracked. Hyphal structure. Hyphal system monomitic; generative hyphae simple-septate; hyphae IKI –, CB –; tissues unchanged in KOH. Subiculum. Generative hyphae hyaline, thin-walled, frequently branched, interwoven, 0.8–1.2 µm in diameter. Hymenium. Cystidia and skeletocystidia not observed. Basidia clavate, with four sterigmata and a simple septum at the base, 15.5–20.1 × 3.5–4.8 µm. Basidioles similar to basidia in shape, but smaller. Spores. Basidiospores broadly ellipsoid to oblong ellipsoid, hyaline, thin-walled, smooth, occasionally with one oil drop, IKI –, CB –, (4.3 –) 5.2–7.9 (– 8.4) × (1.9 –) 2.5–5 (– 5.5) µm, L = 6.44 µm, W = 3.6 µm, Q = 1.51–2.09 (n = 90 / 3). Additional specimens (paratypes) examined. • China; Shaanxi Province, Shangzhou District; 33.761057°N, 109.820994°E; elevation 1187 m; on fallen angiosperm trunk; 22 August 2025; Cui 24843 (BJFC). • China; Shaanxi Province, Shangzhou District; 33.760510°N, 109.821477°E; elevation 1206 m; on fallen angiosperm branch; 22 August 2025; Cui 24842 (BJFC). Type of rot. White rot.

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